Does tool-use necessarily indicate intelligence?
When does tool use evolve?
Is tool behaviour usually learnt?
Does tool-use indicate complex cognition?
Expanding the story
Tool use is ecologically important for Wood-Pecker finches
How does tool use develop in W-P finches?
Tool making: general or specialised cognition?
Social transmission of tool behaviour in Goffin's
Expression of tool use restricted by morphology?
Final word: comparative studies
Tool use as an adaptation
Innovation and Problem-Solving
What is animal problem solving and innovation?
Exploration, behavioural flexibility and neophobia
Consequences for ecology?
A full circle?